Is the DNS info the same for both machines? I know you said that they are both set to pull this info with DHCP, but... are they the same?
Second, is the DNS info being obtained from a server, hosted on site, or through a gateway/router? If the info is coming from a router, then I would suggest logging into the router and obtaining the DNS addresses from there. You will find that they are not private IP addresses.
If DNS is hosted on a server, then check with the IT department and let them know what your running into. They should be able to make some changes to the DNS table to accomidate the MFP's.
Is the SMTP server an in-house server?If so... then I wouldn't sweat the details my friend.
If not.... well, you might want to keep looking at it.
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