I realize this may be a shot in the dark here, but...
Did you try disable cartridge check from the diagnostics menu and switch the cartridges to a different slot? Then try the test again to see if it follows the cartridge or stays with the slot.
Did you try another transfer belt?
I can't think why it would happen when printing but not when copying other than a software/communication issue, but at this point it may not hurt to try these 2 things.
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