This looks like a very unique issue.
At first glance, bias would be my immediate guess. But the negative image on the right side of the page (lead edge) tells me that something else is wrong and that the laser write process has reversed into a negative image.
Just out of curiosity, if you were to copy a good page off the glass and set the machine to give you a negative image as an output, what does the output document look like? If it is the opposite of the images you posted, I would say replace the PRCB/MCB. It might even be memory related.
Bookmarks