Please check if the copier is setup to auto tray selection as well. We usually disable bypass from auto tray selection.
Utility>System setting>Auto Tray Selection Settings] Select all trays→[OK]→[OK]
i'd also check the tray switching setting while you are there
Utility>System setting>Paper tray settings>Autotray switching on
Utility>System setting>Paper tray settings>No matching paper > switch tray
Lastly ill check if the A4/A3>letter/large switching is enabled at utility>Printer settings>Basic/general settings>[A4/A3<-->LTR/LGRAutoSwitchSetting]→[ON]
We always have issues like this due to at least 1 PC at client site setup on different region or they print a email originated from a different region
Hope this helps ...
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